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The Devil Wears Prada star knocked Keira Knightley off the number one spot, due to her acting in films Rachel Getting Married and Becoming Jane, reports Metro.
Editor of LoveFilm.com Helen Cowley said: "This star is seriously on the rise. She has clawed her way to the top of this chart after winning an Oscar nomination for her role in Rachel Getting Married. Our Keira definitely needs to pull her cinematic socks up if she wants to regain the top spot in 2010."
Sienna Miller took tenth place, while Emily Blunt beat Scarlett Johansson and Natalie Portman to be voted fourth best.
Heath Ledger's ex girlfriend Michelle Williams made a surprise entry at number three.
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