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The French actress will star as Syrena, a mermaid who encounters Johnny Depp's Captain Jack Sparrow.
The 23-year-old's previous screen credits include Bruc and La Première Étoile. She has also appeared on stage in the French production of Equus.
Depp, Penélope Cruz, Ian McShane, Geoffrey Rush and Stephen Graham will star in On Stranger Tides, with Rob Marshall and Jerry Bruckheimer serving as director and producer respectively.
Keira Knightley and Orlando Bloom have stepped aside from the fourth nautical blockbuster.
Disney plans to shoot the new Pirates film in June for a May 2011 release.
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