The rockers push Usher and will.i.am down to second place following the pair's two-week reign at the chart summit with 'OMG'.
Glee's cover of Bonnie Tyler classic 'Total Eclipse Of The Heart' climbs up 11 spots to third place, while Alexandra Burke's 'All Night Long' reaches a new peak of fourth.
Jason Derulo's 'Ridin' Solo' is the highest new entry of the week in fifth position. The other main newcomer to the chart is DJ Team's 'Stereo Love' in eighth spot.
The top ten singles in full (click where possible for our reviews):
1. (2) Train: 'Hey, Soul Sister'
2. (1) Usher ft. will.i.am: 'OMG'
3. (14) Glee cast ft. Jonathan Groff: 'Total Eclipse Of The Heart'
4. (16) Alexandra Burke ft. Pitbull: 'All Night Long'
5. (-) Jason Derulo: 'Ridin' Solo'
6. (3) Lady Antebellum: 'Need You Now'
7. (4) Lady GaGa ft. Beyoncé: 'Telephone'
8. (-) DJ Team: 'Stereo Love'
9. (12) Plan B: 'She Said'
10. (7) Timbaland ft. Justin Timberlake: 'Carry Out'