Hamley was described as one of the most-loved animals on the reserve at Glen Afric , where cast including Stephen Tompkinson and Dawn Steele have worked with him for the past five years.
The 7-year-old giraffe died after being struck by lightning during a thunderstorm yesterday.
"We are all desperately upset about the passing of Hamley, who has been part of the Wild At Heart family for five years," producer Nick Goding said. "It was a natural disaster, but nevertheless heartbreaking for everyone who has been involved with him - he was a real character.
"Our wildlife is very much at the heart of the series - Hamley was a gentle, playful and charming animal. He will be greatly missed by everybody".
Wild At Heart returns for its sixth series next year.
> Warren Clarke joins 'Wild At Heart'